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et al., 2016), Phoma Rhizopus , Epicoccum ( 2.2 Factors influencing the community composition of tobacco microorganisms The community composition of tobacco-associated microorganisms is dynamic, and influenced by various environmental factors, such as temperature, humidity and pH ( Pantoea and Variovorax is positively correlated with temperature and humidity, whereas the abundance of Nesterenkonia , Staphylococcus , Chryseomonas , Rhodococcus , Paracoccus , Serratia and Ralstonia shows a negative correlation with temperature and humidity ( Golovinomyces is significantly affected by temperature, while the abundance of Alternaria , Phoma , Trichoderma , Leptosphaerulina , Gibellulopsis and Candida is notably impacted by relative humidity ( Klebsiella variicola , Klebsiella pneumoniae , Serratia , and Salmonella initially increases, peaking after 16 h before subsequently decreasing as the fermentation process continues to 24 and 36 h (Xing et al., 2021)