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iv) Sulfur radical pathways: ROS/RNS generate sulfur radicals (S), leading to disulfide fomation

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However, the low expression levels of recombinant proteins in cultured mammalian cells presents a challenge

(PubMed) Suprapto B, Widardo, Suhanantyo

Crystal structure of the human thioredoxin reductase-thioredoxin complex